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Timo Arula

https://www.etis.ee/CV/Timo_Arula/eng
2.01.1982
443 3800
timo.arula@ut.ee
http://
G-8710-2017
(1) Reviewed manuscripts for following journals (ISI, CC): Journal of Plankton Research (JPR); Marine Ecology Progress Series (MEPS); Estuarine, Coastal and Shelf Science (ECSS); Marine Environmental Research (MERE); Journal of Agricultural Sciences (JAS); Marine Biology Research (MBR); Helgoland Marine Research (HMR), Journal of Oceanology and Limnology
ecology (JOL), Fishes, ICES Journal of Marine Science, Journal of Marine Science and Engineering, Sustainability, Journal of Fish Biology, Journal of Animal Ecology, Marine Policy, Water.

(2) Preparation of lection "Baltis Sea - internationally harvested commercial fish species in the Baltic Sea", course for MSc students.

Career

Institutions and positions
01.01.2021–...   
University of Tartu, Faculty of Science and Technology, Estonian Marine Institute, Associate Professor in Marine Ecosystems (1,00)
01.07.2018–...   
University of Maryland, Center for Environmental Science, Chesapeake Biological Laboratory, Postdoctoral research fellow (1,00)
01.09.2020–31.12.2020   
University of Tartu, Faculty of Science and Technology, Estonian Marine Institute, Senior Research Fellow in Marine Ecosystems (1,00)
01.07.2019–31.08.2020   
University of Tartu, Faculty of Science and Technology, Estonian Marine Institute, Research Fellow in Marine Ecosystems (1,00)
01.01.2016–30.06.2018   
University of Tartu, Faculty of Science and Technology, Estonian Marine Institute, Research Fellow in Marine Ecosystems (1,00)
01.06.2012–31.05.2015   
University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu, Research Fellow in Marine Ecosystems (1,00)
2011–2016   
University of Hamburg, University of Hamburg, The Institute for Hydrobiology and Fisheries Science- several short-term practise. (1,00)
01.01.2008–31.05.2012   
University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu, Graduate student in doctoral studies (1,00)
2006–2006   
University of Southampton, National Oceanography Centre, University of Southampton, National Oceanography Centre - participation in field survey in North Atlantic ocean (1,00)
05.09.2005–31.07.2006   
University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu, graduate student in master studies (1,00)
 
 
R&D related managerial and administrative work
2020−...   
AGRAARTEADUS | JOURNAL OF AGRICULTURAL SCIENCE Editorial Board (Advisory Board)
2017−...   
ICES Working Group of Baltic International Fish Survey (WGBIFS) member
2016−...   
Invited plenary speaker (keynote) and theme session moderator in "Fish biology /early stages". "41th CIESM Congress" 41th Mediterranean Science Conference, Kiel, Germany, 12-16.09.2016
2014−...   
ICES Working Group on Atlantic Fish Larvae and Eggs Surveys (WGALES) member
2013−...   
ICES Working Group of Baltic Fisheries Assessment (WGBFAS) member
2013−...   
Member of Early Life History Section of the American Fisheries Society
2007−...   
Regular participation in annually performed ICES (International Council for Exploration of Sea) coordinated Baltic Sea fish stock surveys - BIAS (Baltic International Acoustical Survey), BASS (Baltic Acoustical Sprat Survey).
 
 
Creative work
Lections on Ecology of the Baltic Sea LOOMI.00.001 (partial reading for master students in University of Tartu). 3,0 EAP
 
 
Additional information
0

Qualifications

Academic degrees
Timo Arula, Doctor's Degree, 2012, (sup) , Ecology of early life-history stages of herring Clupea harengus membras in the northeastern Baltic Sea, University of Tartu.
Timo Arula, Research Master's Degree, 2008, (sup) , Liivi lahe kevadkuduräime Clupea harengus membras vastsete arvukuse dünaamika ja selle seos kujuneva räimepõlvkonna tugevusega (Abundance dynamics of larvae of the spring spawning herring Clupea harengus membras of the Gulf of Riga and its relationship to the year class strength of the stock), Estonian University of Life Sciences, Institute of Agricultural and Environmental Sciences.
 
 
Honours & awards
2017, Timo Arula, Council on lnternational Education Exchange (CIEE), Research Scholar Program award for Postdoctoral Fellowship in University of Maryland, MD, USA.
2014, Timo Arula, European life-long studies project ERASMUS grant to participate in ICES working group
2013, Timo Arula, Scholarship for Estonian summary of non-Estonian doctoral dissertation
2012, Timo Arula, Kristjan Jaak travel grant for young research fellow
2011, Timo Arula, American Fisheries Society, Early Life History Section J.H.S. Blaxter best student conference presentation award from "35th Annual Larval Fish Conference". Wilmington, North-Carolina, USA.
2009, Timo Arula, Estonian Students Fund grant for PhD studies
2008, Timo Arula, ESF DoRa progam for Supporting the participation of young researchers in international exchange of knowledge
2007, Timo Arula, Estonian World Council grant for PhD studies
 
 
Fields of research
ETIS CLASSIFICATION: 1. Biosciences and Environment; 1.4. Ecology, Biosystematics and -physiology; CERCS CLASSIFICATION: B260 Hydrobiology, marine biology, aquatic ecology, limnology
 
 
Additional information
Doctor of Philosophy
University of Tartu, 2012
Invited speaker (title: Over a half century of larval herring studies in the Gulf of Riga (Baltic Sea) - what we have learned?) GEOMAR (Helmholtz Centre for Ocean Research) scientific seminar. Kiel, Germany, 1.12.2016
Invited speaker (title: Over a half century of larval herring studies in the Gulf of Riga (Baltic Sea) - what we have learned?) at UHAM (Universität Hamburg, Institut für Hydrobiologie und Fischereiwissenschaft) scientific seminar. Hamburg, Germany, 2.12.2016
Distinguished Scholar Seminar speaker (title:Seventy years of fish early life history studies in NE Baltic Sea: What we have learned?) UMCES CBL (Chesapeake Biological Laboratory). Maryland, USA. 31.10.2018
Seminar speaker (title: How do we best determine a bad recruitment year?) IMR (Institute of Marine Research). Bergen, Norway. 17.01.2020
Seminar speaker (title: Seventy years of larval herring studies in the Gulf of Riga (Baltic Sea) to improve our understanding on recruitment processes) IMR (Institute of Marine Research). Flödevigen, Norway. 17.01.2020

Projects in progress

Completed projects

Dissertations under supervision

Supervised dissertations

Publications

1.1. Scholarly articles indexed by Web of Science Science Citation Index Expanded, Social Sciences Citation Index, Arts & Humanities Citation Index, Emerging Sources Citation Index and/or indexed by Scopus (excluding chapters in books)
 
Auth, T.; Arula, T.; Houde, D., E.; Woodland, J., R. (2020). Spatial ecology and growth in early-life stages of bay anchovy (Anchoa mitchilli) in Chesapeake Bay (USA). Marine Ecology Progress Series, 651, 125−143. DOI: 10.3354/meps13431.
 
Liversage, K.; Kotta, J.; Aps, R.; Fetissov, M.; Nurkse, K.; Orav-Kotta, H.; Rätsep, M.; Forsström, T.; Fowler, A.; Lehtiniemi, M.; Normant-Saremba, M.; Puntila-Dodd, R.; Arula, T.; Hubel, K.; Ojaveer, H. (2019). Knowledge to decision in dynamic seas: Methods to incorporate non-indigenous species into cumulative impact assessments for Maritime Spatial Planning. The Science of The Total Environment, 658, 1452−1464. DOI: 10.1016/j.scitotenv.2018.12.123.
 
Arula, Timo; Shpilev, Heli; Raid, Tiit; Sepp, Elor (2019). Thermal conditions and age structure determine the spawning regularities and condition of Baltic herring (Clupea harengus membras) in the NE of the Baltic Sea. PeerJ, 7, ARTN e7345.10.7717/peerj.7345.
 
Patrik Kraufvelin, Zeynep Pekcan-Hekim, Ulf Bergström, Ann-Britt Florin, Annukka Lehikoinen, Johanna Mattila, Timo Arula, Laura Briekmane, Elliott John Brown, Zuzanna Celmer, Justas Dainys, Henri Jokinen, Petra Kääriä, Meri Kallasvuo, Antti Lappalainen, Linas Lozys, Peter Möller, Alessandro Orio, Mehis Rohtla, Lauri Saks, Martin Snickars, Josianne Støttrup, Göran Sundblad, Imre Taal, Didzis Ustups, Aare Verliin, Markus Vetemaa, Helmut Winkler, Adam Wozniczka, Jens Olsson (2018). Essential coastal habitats for fish in the Baltic Sea. Estuarine Coastal and Shelf Science, 204, 14−30.10.1016/j.ecss.2018.02.014.
 
Kotta, J.; Herkül, K.; Jaagus, J.; Kaasik, A.; Raudsepp, U.; Alari, V.; Arula, T.; Haberman, J.; Järvet, A.; Kangur, K.; Kont, A.; Kull, A.; Laanemets, J.; Maljutenko, I.; Männik, A.; Nõges, P.; Nõges, T.; Ojaveer, H.; Peterson, A.; Reihan, A. ... Tõnisson, H. (2018). Linking atmospheric, terrestrial and aquatic environments in high latitude: Regime shifts in the Estonian regional climate system for the past 50 years. PLoS ONE, 13 (12), ARTN e0209568−20. DOI: 10.1371/journal.pone.0209568.
 
Arula, Timo; Shpilev, Heli; Raid, Tiit; Vetemaa, Markus; Albert, Anu (2017). Maturation at a young age and small size of European smelt (Osmerus eperlanus): A consequence of population overexploitation or climate change? Helgoland Marine Research, 71, ARTN 7. DOI: 10.1186/s10152-017-0487-x.
 
Ojaveer, H.; Gross, R.; Laur, K.; Arula, T.; Klais, R. (2017). Genetic analysis reveals the diversity of larval Gobiidae in a temperate estuary. Journal of Fish Biology, 91 (4), 1048−1061.10.1111/jfb.13407.
 
Arula, T.; Raid, T.; Simm, M.; Ojaveer, H. (2016). Temperature-driven changes in early life-history stages influence the Gulf of Riga spring spawning herring (Clupea harengus m.) recruitment abundance. Hydrobiologia, 767 (1), 125−135. DOI: 10.1007/s10750-015-2486-8.
 
Bekkevold, Dorte; Gross, Riho; Arula, Timo; Helyar, Sarah J; Ojaveer, Henn (2016). Outlier loci detect intraspecific biodiversity amongst spring and autumn spawning herring across local scales. PLoS ONE, 11 (4, e0148499), 1−13. DOI: 10.1371/journal.pone.0148499.
 
Ojaveer, H.; Tomkiewicz, J.; Arula, T.; Klais, R. (2015). Female ovarian abnormalities and reproductive failure of autumn-spawning herring (Clupea harengus membras) in the Baltic Sea. ICES Journal of Marine Science, 72 (8), 2332−2340.10.1093/icesjms/fsv103.
 
Arula, T; Laur, K.; Simm, M.; Ojaveer, H. (2015). Dual impact of temperature on growth and mortality of marine fish larvae in a shallow estuarine habitat. Estuarine Coastal and Shelf Science, 167, 326−335. DOI: 10.1016/j.ecss.2015.10.004.
 
Arula, T.; Ojaveer, H.; Klais, R. (2014). Impact of extreme climate and bioinvasion on the temporal coupling of spring herring (Clupea harengus m.) larvae and their prey. Marine Environmental Research, 102, 102−109.10.1016/j.marenvres.2014.05.001.
 
Arula, T.; Gröger, J.; Ojaveer, H.; Simm, M. (2014). Shifts in the spring herring (Clupea harengus membras) larvae and related environment in the eastern Baltic Sea over the past 50 years. PLoS ONE, 9 (3), e91304.10.1371/journal.pone.0091304.
 
Müller-Karulis, B.; Arula, T.; Balode, M.; Laur, K.; Ojaveer, E. (2013). Challenges and opportunities of local fisheries management: pikeperch, Sander lucioperca (Actinopterygii: Perciformes: Percidae), in Pärnu Bay, northern Gulf of Riga, Baltic Sea. Acta Ichthyologica Et Piscatoria, 43 (2), 151−161.10.3750/AIP2013.43.2.08.
 
Arula, T.; Kotta, J.; Lankov, A.; Simm, M.; Põlme, S. (2012). Diet composition and feeding activity of larval spring-spawning herring: importance of environmental variability. Journal of Sea Research, 68, 33−40. DOI: 10.1016/j.seares.2011.12.003.
 
Arula, T.; Ojaveer, H.; Shpilev, H. (2012). Individual fecundity of the autumn spawning Baltic herring Clupea harengus membras L. Estonian Journal of Ecology, 61 (2), 119−134.10.3176/eco.2012.2.04.
 
Ojaveer, Evald; Arula, Timo; Lankov, Ain; Shpilev, Heli (2011). Impact of environmental deviations on the larval and year class abundances in the spring spawning herring (Clupea harengus membras L.) of the Gulf of Riga (Baltic Sea) in 1947 -- 2004. Fisheries Research, 107, 159−168.
2.3. Dissertations published in a series of dissertations (excluding manuscripts)
 
Arula, T. (2012). Ecology of early life-history stages of herring Clupea harengus membras in the northeastern Baltic Sea. (Tartu Ülikool). Tartu University Press.
2.5. Published research project report or study
 
Raudsepp, Urmas; Jaagus, Jaak; Alari, Victor; Arula, Timo; Järvet, Arvo; Kont, Are; Kotta, Jonne; Kull, Ain; Laanemets, Jaan; Maljutenko, Ilja; Männik, Aarne; Reihan, Alvina; Rõõm, Rein; Sepp, Mait; Suursaar, Ülo; Tamm, Ottar; Tamm, Toomas; Tõnisson, Hannes (2015). Eesti kliima ja keskkonnaseisundi võimalike muutuste hindamine atmosfääri-, mere- ja jõgede äravoolu dünaamiliste mudelite tulemuste põhjal (ESTKLIIMA). Andres Tõnisson (Toim.). Keskkonnakaitse ja -tehnoloogia teadus- ja arendustegevuse programmi teaberaamat 2010-2015 (88−94). Eesti Teadusagentuur.
3.1. Articles/chapters in books published by the publishers listed in Annex (including collections indexed by the Web of Science Book Citation Index, Web of Science Conference Proceedings Citation Index, Scopus)
 
Sepp, Elor; Raid, Tiit; Arula, Timo (2020). Does spatial patterns in fishing explain dynamics of commercial pelagic populations in Baltic Sea? Taylor & Francis Group, 617−621. DOI: 10.1201/9780367810085-82.
 
Raid, Tiit; Arula, Timo; Kaljuste, Olavi; Sepp, Elor; Järv, Leili; Hallang, Andrus; Shpilev, Heli; Lankov, Ain (2015). Dynamics of the commercial fishery in the Baltic Sea: what are the driving forces? Towards Green Marine Technology and Transport: Proceeding of the 16th International Congress of the International Maritime Assotiation of the Mediterranean (IMAM 2015), Pula, Croatia, 21-24 September. Ed. C. Guedes Soares; R. Dejhalla; D. Pavletic´. Taylor & Francis, 897−906.
3.4. Articles/presentations published in conference proceedings not listed in Section 3.1
 
Arula, Timo (2016). CIESM Congress Session : Fish biology / early stages (Moderator´s Synthesis). In: Silvia Ceramicola, Miroslav Gačić, François Galgani, Frank Oliver Glöckner, Jamila Ben Souissi, Salud Deudero, Tamara Shiganova, Yves Henocque (_EditorsAbbr). 41st CIESM Congress Proceedings (329−329). Monaco: Rapp. Comm. int. Mer Médit. (41).
5.2. Conference abstracts that do not belong to section 5.1
 
Arula, T; Woodland J. Ryan; Houde D. Edward (2019). DEFINING THE TROPHIC ROLE OF A KEY FORAGE SPECIES IN A LARGE COASTAL ECOSYSTEM. San Juan, Puerto Rico. 24.02-1.03.2019.
 
Arula, T.; Raid, T.; Sepp, E.; Shpilev, H. (2017). The spawning, age structure and condition of Baltic spring spawning herring (Clupea harengus membras) is significantly modified by winter conditions prior spawning. 1st Southeast European Ichthyological Conference (SEEIC 2017). 27-29.09.2017 University of Sarajevo.
 
Arula, Timo (2016). Keynote in theme session "Fish biology / early stages". In: 41th Mediterranean Science Commission Congress, CIESM.41th CIESM Congress (Mediterranean Sea Science Congress). Christian Albrechts University, Kiel.
 
Arula, T.; Raid, T.; Simm, M.; Ojaveer, H. (2015). Factors affecting the abundance of spring spawning herring (Clupea harengus m.) larvae in the Gulf of Riga.
 
Arula, T.; Shpilev, H. (2015). Early ontogenesis of European smelt (Osmerus eperlanus) in variable environments.
 
Arula, T.; Ojaveer, H.; Raid, T.; Simm, M. (2015). Importance of plankton seasonality on larval herring and year-class abundance of the Gulf of Riga spring spawning herring (Clupea harengus m.).
 
Arula, T.; Ojaveer, H.; Raid, T. (2015). Mortality and growth at larval stage: advancing the understanding of stock dynamics processes in the Gulf of Riga spring spawning herring (Clupea harengus membras).
 
Arula, T.; Shpilev, H.; Raid, T. (2014). Discrimination of herring ecotypes using otolith shape and body morphology in the Baltic Sea. 165−165.
 
Arula, T.; Laur, K.; Ojaveer, H. (2014). Small-scale dynamics of vital rates of herring Clupea harengus membras and goby Pomatoschistus spp. larvae in a shallow temperate bay. Johan Hjort Symposium on Recruitment Dynamics and Stock Variability, Bergen, Norway, 7- 9 October 2014. Norway,.
 
Arula, T.; Gröger, J.; Ojaveer, H.; Simm, M. (2013). Indication for a regime shift in the larval spring herring (Clupea harengus membras) and the related environment in the Gulf of Riga (Baltic Sea). 37th Annual larval Fish Conference, Miami, USA. The Rosenstiel School of Marine and Atmospheric Science (RSMAS) of the University of Miami. 2-6.06.. http://www.rsmas.miami.edu/LFC2013/LFC2013program.pdf,.
 
Arula, T; Ojaveer, H; Klais, R (2013). Impact of extreme climate and bioinvasion on phenology of spring herring (Clupea harengus m.) larvae and their prey.
 
Arula, T; Ojaveer, H; Klais, R. (2013). Multidecadal abundance and phenology dynamics of zooplankton and fish larvae in a shallow temparate bay.
 
Arula, Timo; Gröger, Paul, Joachim; Ojaveer, Henn; Simm, Mart (2012). Application of shiftograms to track abrupt changes in marine ecosystems: case study in the Gulf of Riga (Baltic Sea) for 1957-2010.
 
Arula, T.; Ojaveer, H. (2012). Life on the edge: past and present of the autumn spawning herring in the Gulf of Riga (Baltic Sea).
 
Arula, T.; Simm, M.; Põllupüü, M.; Ojaveer, H. (2011). LONG TERM CHANGES IN PHENOLOGY OF COPEPODS AND HERRING LARVAE IN THE GULF OF RIGA (BALTIC SEA).
 
Arula, T.; Kotta, J.; Lankov, A.; Simm, M.; Põlme, S. (2011). Diet composition and feeding activity of larval spring-spawning herring: importance of environmental variability. 35th Annual Larval Fish Conference; Wilmington, North Carolina, USA; May 22-25. Ed. Fred Scharf; Jeff Buckel; Tom Lankford., x−x.
 
Arula, T.; Shpilev, H.; Ojaveer, H. (2009). Fecundity of the autumn spawning herring in the Gulf of Riga (Baltic Sea): comparison between 1950s-1970s and 2000s. BSSC 2009 ABSTRACT BOOK: 7th Baltic Sea Science Congreess; Tallinn, Estonia; August 17-21, 2009. Ed. Lips, U. Tallinn University of Technology: Infotrükk OÜ, 108−108.
 
Arula, T.; Ojaveer, E.; Ojaveer, H.; Põllupüü, M. (2008). LONG-TERM DYNAMICS OF THE SPRING-SPAWNING HERRING LARVAE (CLUPEA HARENGUS MEMBRAS) IN THE GULF OF RIGA (BALTIC SEA). 32nd Annual Larval Fish Conference: 32nd Annual Larval Fish Conference; Kiel, Germany; 4-7 August 2008.. Ed. Catriona Clemmesen. Germany, Kiel, 56−56.
1.02.2021

Timo Arula

https://www.etis.ee/CV/Timo_Arula/est
2.01.1982
443 3800
timo.arula@ut.ee
G-8710-2017
(1) Retsenseerinud artiklite käsikirju rahvusvahelistes eelretsentseeritavates ajakirjades (ISI, CC): Journal of Plankton Research; Marine Ecology Progress Series; Estuarine, Coastal and Shelf Science; Marine Environmental Research; Journal of Agricultural Sciences; Marine Biology Research; Helgoland Marine Research, Journal of Oceanology and Limnology, Fishes, ICES Journal of Marine Science, Journal of Marine Science and Engineering, Sustainability, Journal of Fish Biology, Journal of Animal Ecology, Marine Policy, Water.

(2) Loengumaterjalide koostamine Tartu Ülikooli magistriõppekava aine "Läänemere ökoloogia" Läänemere rahvusvaheliselt majandatavate töönduskalade ökoloogia osale 4*45 minutit.

Teenistuskäik

Töökohad ja ametid
01.01.2021–...   
Tartu Ülikool, Loodus- ja täppisteaduste valdkond, Eesti mereinstituut, meresüsteemide kaasprofessor (1,00)
01.07.2018–...   
Marylandi Ülikool, Keskkonnateaduste Keskus, Chesapeake Bioloogia Labor, Järeldoktor (1,00)
01.09.2020–31.12.2020   
Tartu Ülikool, Loodus- ja täppisteaduste valdkond, Eesti mereinstituut, meresüsteemide vanemteadur (1,00)
01.07.2019–31.08.2020   
Tartu Ülikool, Loodus- ja täppisteaduste valdkond, Eesti mereinstituut, meresüsteemide teadur (1,00)
01.01.2016–30.06.2018   
Tartu Ülikool, Loodus- ja täppisteaduste valdkond, Eesti mereinstituut, meresüsteemide teadur (1,00)
01.06.2012–31.05.2015   
Tartu Ülikool, Loodus- ja tehnoloogiateaduskond, Tartu Ülikooli Eesti Mereinstituut, mereökosüsteemide teadur (1,00)
2011–2016   
University of Hamburg, Hamburgi Ülikool, Hüdrobioloogia ja kalandusteaduste Instituut- mitmed episoodilised stažeerimised (1,00)
01.01.2008–31.05.2012   
Tartu Ülikool, Loodus- ja tehnoloogiateaduskond, Tartu Ülikooli Eesti Mereinstituut, kraadiõppur doktorantuuris (1,00)
2006–2006   
University of Southampton, National Oceanography Centre, Southamptoni Ülikool, Riiklik Okenograafia Keskus - osalemine uurimislaeva "DISCOVERY" ekspeditsioonil Põhja-Atlandi ookeanil (1,00)
05.09.2005–31.07.2006   
Tartu Ülikool, Loodus- ja tehnoloogiateaduskond, Tartu Ülikooli Eesti Mereinstituut, kraadiõppur magistrantuuris (1,00)
 
 
Teadusorganisatsiooniline ja -administratiivne tegevus
2020−...   
AGRAARTEADUS | JOURNAL OF AGRICULTURAL SCIENCE Toimetuskolleegiumi liige (Advisiory Board)
2017−...   
Rahvusvahelise Mereuurimisnõukogu (ICES) Läänemere rahvusvaheliselt majandatavate kalavarude uuringute töörühma (WGBIFS) liige
2016−...   
41-l Vahemere Teaduskonverentsil "41th CIESM Congress" teemasesiooni "Fish Biology/ early stages" juhatamine ja sissejuhatava plenaar ettekande pidamine. Kiel, Saksamaa, 12-16.09.2016
2014−...   
Rahvusvahelise Mereuurimisnõukogu (ICES) Atlandi ookeani kalavastsete ja -marja uuringute töörühma (WGALES) liige
2013−...   
Rahvusvahelise Mereuurimisnõukogu (ICES) Läänemere kalavarude suuruse hindamise töörühma (WGBFAS) liige
2013−...   
Ameerika Kalandusühingu Kalade Varajaste Elustaadiumite seltsi liige (Early Life History Section of the American Fisheries Society)
2007−...   
Regulaarne osalemine iga-aastaselt toimuvatel ICES (Rahvusvaheline Mereuurimise Organisatsioon) koordineeritavate Läänemere kalavarude uuringutel - BIAS (Baltic International Acoustical Survey), BASS (Baltic Acoustical Sprat Survey).
 
 
Loometöö
Bioloogia, Keskkonnatehnoloogia, Ökoloogia ning elustiku kaitse eriala (Tartu Ülikooli magistriõppekava), Läänemere ökoloogia loeng LOOMI.00.001 (osaline läbiviimine), Maht 3,0 EAP.


 
 
Lisainfo
0

Kvalifikatsioon

Teaduskraadid
Timo Arula, Doctor's Degree, 2012, (sup) , Ecology of early life-history stages of herring Clupea harengus membras in the northeastern Baltic Sea, University of Tartu.
Timo Arula, Research Master's Degree, 2008, (sup) , Liivi lahe kevadkuduräime Clupea harengus membras vastsete arvukuse dünaamika ja selle seos kujuneva räimepõlvkonna tugevusega (Abundance dynamics of larvae of the spring spawning herring Clupea harengus membras of the Gulf of Riga and its relationship to the year class strength of the stock), Estonian University of Life Sciences, Institute of Agricultural and Environmental Sciences.
 
 
Teaduspreemiad ja tunnustused
2017, Timo Arula, Rahvusvahelise Haridusvahetuse Nõukogu (CIEE) teadustöö stipendium järeldoktoriks Marylandi Ülikooli, Ameerika Ühendriigid.
2014, Timo Arula, Euroopa elukestva õppe progammi ERASMUS stipendium osalemaks Rahvusvahelise mereuurimisnõukogu töörühmal
2013, Timo Arula, Võõrkeelse väitekirja eestikeelse resümee stipendiumi Tartu Ülikooli sihtasutuselt programmi „Eestikeelse terminoloogia toetamine" raames
2012, Timo Arula, Kristjan Jaagu välisõidu stipendium noorele teadlasele
2011, Timo Arula, Kalade Varajaste Elustaadiumite Selts USAs, J.H.S. Blaxter ´i-nimeline parima teaduskonverentsi ettekande auhind "35lt Iga-aastaselt Kalade Varajaste Elustaadiumite Konverentsilt". Wilmington, North-Carolina, USA.
2009, Timo Arula, Eesti Üliõpilaste Toetusfondi USAs stipendium doktoriõpinguteks
2008, Timo Arula, ESF progammi DoRa noorteadlase stipendium osalemaks rahvusvahelises teadmisteringluses
2007, Timo Arula, Ülemaailmse Eesti Kesknõukogu Margot M. ja Herbert R. Linna stipendium doktoriõpinguteks
 
 
Teadustöö põhisuunad
ETIS KLASSIFIKAATOR: 1. Bio- ja keskkonnateadused; 1.4. Ökoloogia, biosüstemaatika ja -füsioloogia; CERCS KLASSIFIKAATOR: B260 Hüdrobioloogia, mere-bioloogia, veeökoloogia, limnoloogia
 
 
Lisainfo
teadusdoktor
Tartu Ülikool, 2012
Külalisloeng (60 minutit) teemal "Over a half century of larval herring studies in the Gulf of Riga (Baltic Sea) - what we have learned?" (Helmholtz Centre for Ocean Research) teadusseminaril. Kiil, Saksamaa, 1.12.2016
Külalisloeng (60 minutit) teadusseminaril teemal "Over a half century of larval herring studies in the Gulf of Riga (Baltic Sea) - what we have learned?" UHAM (Universität Hamburg, Institut für Hydrobiologie und Fischereiwissenschaft) teadusseminaril. Hamburg, Saksamaa, 2.12.2016
Külalisloeng (60 minutit) teemal "Seventy years of fish early life history studies in NE Baltic Sea: What we have learned?" CBL (Chesapeake Biological Laboratory) teadusseminaril. Maryland, USA. 31.10.2018
Külalisloeng (60 minutit) teemal "How do we best determine a bad recruitment year?" IMR (Institute of Marine Research) teadusseminaril. Bergen, Norra. 16.01.2020
Külalisloeng (60 minutit) teemal "Seventy years of larval herring studies in the Gulf of Riga (Baltic Sea) to improve our understanding on recruitment processes?" IMR (Institute of Marine Research) teadusseminaril. Flödevigen, Norra. 17.01.2020

Jooksvad projektid

Lõppenud projektid

Juhendamisel väitekirjad

Juhendatud väitekirjad

Publikatsioonid

1.1. Scholarly articles indexed by Web of Science Science Citation Index Expanded, Social Sciences Citation Index, Arts & Humanities Citation Index, Emerging Sources Citation Index and/or indexed by Scopus (excluding chapters in books)
 
Auth, T.; Arula, T.; Houde, D., E.; Woodland, J., R. (2020). Spatial ecology and growth in early-life stages of bay anchovy (Anchoa mitchilli) in Chesapeake Bay (USA). Marine Ecology Progress Series, 651, 125−143. DOI: 10.3354/meps13431.
 
Liversage, K.; Kotta, J.; Aps, R.; Fetissov, M.; Nurkse, K.; Orav-Kotta, H.; Rätsep, M.; Forsström, T.; Fowler, A.; Lehtiniemi, M.; Normant-Saremba, M.; Puntila-Dodd, R.; Arula, T.; Hubel, K.; Ojaveer, H. (2019). Knowledge to decision in dynamic seas: Methods to incorporate non-indigenous species into cumulative impact assessments for Maritime Spatial Planning. The Science of The Total Environment, 658, 1452−1464. DOI: 10.1016/j.scitotenv.2018.12.123.
 
Arula, Timo; Shpilev, Heli; Raid, Tiit; Sepp, Elor (2019). Thermal conditions and age structure determine the spawning regularities and condition of Baltic herring (Clupea harengus membras) in the NE of the Baltic Sea. PeerJ, 7, ARTN e7345.10.7717/peerj.7345.
 
Patrik Kraufvelin, Zeynep Pekcan-Hekim, Ulf Bergström, Ann-Britt Florin, Annukka Lehikoinen, Johanna Mattila, Timo Arula, Laura Briekmane, Elliott John Brown, Zuzanna Celmer, Justas Dainys, Henri Jokinen, Petra Kääriä, Meri Kallasvuo, Antti Lappalainen, Linas Lozys, Peter Möller, Alessandro Orio, Mehis Rohtla, Lauri Saks, Martin Snickars, Josianne Støttrup, Göran Sundblad, Imre Taal, Didzis Ustups, Aare Verliin, Markus Vetemaa, Helmut Winkler, Adam Wozniczka, Jens Olsson (2018). Essential coastal habitats for fish in the Baltic Sea. Estuarine Coastal and Shelf Science, 204, 14−30.10.1016/j.ecss.2018.02.014.
 
Kotta, J.; Herkül, K.; Jaagus, J.; Kaasik, A.; Raudsepp, U.; Alari, V.; Arula, T.; Haberman, J.; Järvet, A.; Kangur, K.; Kont, A.; Kull, A.; Laanemets, J.; Maljutenko, I.; Männik, A.; Nõges, P.; Nõges, T.; Ojaveer, H.; Peterson, A.; Reihan, A. ... Tõnisson, H. (2018). Linking atmospheric, terrestrial and aquatic environments in high latitude: Regime shifts in the Estonian regional climate system for the past 50 years. PLoS ONE, 13 (12), ARTN e0209568−20. DOI: 10.1371/journal.pone.0209568.
 
Arula, Timo; Shpilev, Heli; Raid, Tiit; Vetemaa, Markus; Albert, Anu (2017). Maturation at a young age and small size of European smelt (Osmerus eperlanus): A consequence of population overexploitation or climate change? Helgoland Marine Research, 71, ARTN 7. DOI: 10.1186/s10152-017-0487-x.
 
Ojaveer, H.; Gross, R.; Laur, K.; Arula, T.; Klais, R. (2017). Genetic analysis reveals the diversity of larval Gobiidae in a temperate estuary. Journal of Fish Biology, 91 (4), 1048−1061.10.1111/jfb.13407.
 
Arula, T.; Raid, T.; Simm, M.; Ojaveer, H. (2016). Temperature-driven changes in early life-history stages influence the Gulf of Riga spring spawning herring (Clupea harengus m.) recruitment abundance. Hydrobiologia, 767 (1), 125−135. DOI: 10.1007/s10750-015-2486-8.
 
Bekkevold, Dorte; Gross, Riho; Arula, Timo; Helyar, Sarah J; Ojaveer, Henn (2016). Outlier loci detect intraspecific biodiversity amongst spring and autumn spawning herring across local scales. PLoS ONE, 11 (4, e0148499), 1−13. DOI: 10.1371/journal.pone.0148499.
 
Ojaveer, H.; Tomkiewicz, J.; Arula, T.; Klais, R. (2015). Female ovarian abnormalities and reproductive failure of autumn-spawning herring (Clupea harengus membras) in the Baltic Sea. ICES Journal of Marine Science, 72 (8), 2332−2340.10.1093/icesjms/fsv103.
 
Arula, T; Laur, K.; Simm, M.; Ojaveer, H. (2015). Dual impact of temperature on growth and mortality of marine fish larvae in a shallow estuarine habitat. Estuarine Coastal and Shelf Science, 167, 326−335. DOI: 10.1016/j.ecss.2015.10.004.
 
Arula, T.; Ojaveer, H.; Klais, R. (2014). Impact of extreme climate and bioinvasion on the temporal coupling of spring herring (Clupea harengus m.) larvae and their prey. Marine Environmental Research, 102, 102−109.10.1016/j.marenvres.2014.05.001.
 
Arula, T.; Gröger, J.; Ojaveer, H.; Simm, M. (2014). Shifts in the spring herring (Clupea harengus membras) larvae and related environment in the eastern Baltic Sea over the past 50 years. PLoS ONE, 9 (3), e91304.10.1371/journal.pone.0091304.
 
Müller-Karulis, B.; Arula, T.; Balode, M.; Laur, K.; Ojaveer, E. (2013). Challenges and opportunities of local fisheries management: pikeperch, Sander lucioperca (Actinopterygii: Perciformes: Percidae), in Pärnu Bay, northern Gulf of Riga, Baltic Sea. Acta Ichthyologica Et Piscatoria, 43 (2), 151−161.10.3750/AIP2013.43.2.08.
 
Arula, T.; Kotta, J.; Lankov, A.; Simm, M.; Põlme, S. (2012). Diet composition and feeding activity of larval spring-spawning herring: importance of environmental variability. Journal of Sea Research, 68, 33−40. DOI: 10.1016/j.seares.2011.12.003.
 
Arula, T.; Ojaveer, H.; Shpilev, H. (2012). Individual fecundity of the autumn spawning Baltic herring Clupea harengus membras L. Estonian Journal of Ecology, 61 (2), 119−134.10.3176/eco.2012.2.04.
 
Ojaveer, Evald; Arula, Timo; Lankov, Ain; Shpilev, Heli (2011). Impact of environmental deviations on the larval and year class abundances in the spring spawning herring (Clupea harengus membras L.) of the Gulf of Riga (Baltic Sea) in 1947 -- 2004. Fisheries Research, 107, 159−168.
2.3. Dissertations published in a series of dissertations (excluding manuscripts)
 
Arula, T. (2012). Ecology of early life-history stages of herring Clupea harengus membras in the northeastern Baltic Sea. (Tartu Ülikool). Tartu University Press.
2.5. Published research project report or study
 
Raudsepp, Urmas; Jaagus, Jaak; Alari, Victor; Arula, Timo; Järvet, Arvo; Kont, Are; Kotta, Jonne; Kull, Ain; Laanemets, Jaan; Maljutenko, Ilja; Männik, Aarne; Reihan, Alvina; Rõõm, Rein; Sepp, Mait; Suursaar, Ülo; Tamm, Ottar; Tamm, Toomas; Tõnisson, Hannes (2015). Eesti kliima ja keskkonnaseisundi võimalike muutuste hindamine atmosfääri-, mere- ja jõgede äravoolu dünaamiliste mudelite tulemuste põhjal (ESTKLIIMA). Andres Tõnisson (Toim.). Keskkonnakaitse ja -tehnoloogia teadus- ja arendustegevuse programmi teaberaamat 2010-2015 (88−94). Eesti Teadusagentuur.
3.1. Articles/chapters in books published by the publishers listed in Annex (including collections indexed by the Web of Science Book Citation Index, Web of Science Conference Proceedings Citation Index, Scopus)
 
Sepp, Elor; Raid, Tiit; Arula, Timo (2020). Does spatial patterns in fishing explain dynamics of commercial pelagic populations in Baltic Sea? Taylor & Francis Group, 617−621. DOI: 10.1201/9780367810085-82.
 
Raid, Tiit; Arula, Timo; Kaljuste, Olavi; Sepp, Elor; Järv, Leili; Hallang, Andrus; Shpilev, Heli; Lankov, Ain (2015). Dynamics of the commercial fishery in the Baltic Sea: what are the driving forces? Towards Green Marine Technology and Transport: Proceeding of the 16th International Congress of the International Maritime Assotiation of the Mediterranean (IMAM 2015), Pula, Croatia, 21-24 September. Ed. C. Guedes Soares; R. Dejhalla; D. Pavletic´. Taylor & Francis, 897−906.
3.4. Articles/presentations published in conference proceedings not listed in Section 3.1
 
Arula, Timo (2016). CIESM Congress Session : Fish biology / early stages (Moderator´s Synthesis). In: Silvia Ceramicola, Miroslav Gačić, François Galgani, Frank Oliver Glöckner, Jamila Ben Souissi, Salud Deudero, Tamara Shiganova, Yves Henocque (_EditorsAbbr). 41st CIESM Congress Proceedings (329−329). Monaco: Rapp. Comm. int. Mer Médit. (41).
5.2. Conference abstracts that do not belong to section 5.1
 
Arula, T; Woodland J. Ryan; Houde D. Edward (2019). DEFINING THE TROPHIC ROLE OF A KEY FORAGE SPECIES IN A LARGE COASTAL ECOSYSTEM. San Juan, Puerto Rico. 24.02-1.03.2019.
 
Arula, T.; Raid, T.; Sepp, E.; Shpilev, H. (2017). The spawning, age structure and condition of Baltic spring spawning herring (Clupea harengus membras) is significantly modified by winter conditions prior spawning. 1st Southeast European Ichthyological Conference (SEEIC 2017). 27-29.09.2017 University of Sarajevo.
 
Arula, Timo (2016). Keynote in theme session "Fish biology / early stages". In: 41th Mediterranean Science Commission Congress, CIESM.41th CIESM Congress (Mediterranean Sea Science Congress). Christian Albrechts University, Kiel.
 
Arula, T.; Raid, T.; Simm, M.; Ojaveer, H. (2015). Factors affecting the abundance of spring spawning herring (Clupea harengus m.) larvae in the Gulf of Riga.
 
Arula, T.; Shpilev, H. (2015). Early ontogenesis of European smelt (Osmerus eperlanus) in variable environments.
 
Arula, T.; Ojaveer, H.; Raid, T.; Simm, M. (2015). Importance of plankton seasonality on larval herring and year-class abundance of the Gulf of Riga spring spawning herring (Clupea harengus m.).
 
Arula, T.; Ojaveer, H.; Raid, T. (2015). Mortality and growth at larval stage: advancing the understanding of stock dynamics processes in the Gulf of Riga spring spawning herring (Clupea harengus membras).
 
Arula, T.; Shpilev, H.; Raid, T. (2014). Discrimination of herring ecotypes using otolith shape and body morphology in the Baltic Sea. 165−165.
 
Arula, T.; Laur, K.; Ojaveer, H. (2014). Small-scale dynamics of vital rates of herring Clupea harengus membras and goby Pomatoschistus spp. larvae in a shallow temperate bay. Johan Hjort Symposium on Recruitment Dynamics and Stock Variability, Bergen, Norway, 7- 9 October 2014. Norway,.
 
Arula, T.; Gröger, J.; Ojaveer, H.; Simm, M. (2013). Indication for a regime shift in the larval spring herring (Clupea harengus membras) and the related environment in the Gulf of Riga (Baltic Sea). 37th Annual larval Fish Conference, Miami, USA. The Rosenstiel School of Marine and Atmospheric Science (RSMAS) of the University of Miami. 2-6.06.. http://www.rsmas.miami.edu/LFC2013/LFC2013program.pdf,.
 
Arula, T; Ojaveer, H; Klais, R (2013). Impact of extreme climate and bioinvasion on phenology of spring herring (Clupea harengus m.) larvae and their prey.
 
Arula, T; Ojaveer, H; Klais, R. (2013). Multidecadal abundance and phenology dynamics of zooplankton and fish larvae in a shallow temparate bay.
 
Arula, Timo; Gröger, Paul, Joachim; Ojaveer, Henn; Simm, Mart (2012). Application of shiftograms to track abrupt changes in marine ecosystems: case study in the Gulf of Riga (Baltic Sea) for 1957-2010.
 
Arula, T.; Ojaveer, H. (2012). Life on the edge: past and present of the autumn spawning herring in the Gulf of Riga (Baltic Sea).
 
Arula, T.; Simm, M.; Põllupüü, M.; Ojaveer, H. (2011). LONG TERM CHANGES IN PHENOLOGY OF COPEPODS AND HERRING LARVAE IN THE GULF OF RIGA (BALTIC SEA).
 
Arula, T.; Kotta, J.; Lankov, A.; Simm, M.; Põlme, S. (2011). Diet composition and feeding activity of larval spring-spawning herring: importance of environmental variability. 35th Annual Larval Fish Conference; Wilmington, North Carolina, USA; May 22-25. Ed. Fred Scharf; Jeff Buckel; Tom Lankford., x−x.
 
Arula, T.; Shpilev, H.; Ojaveer, H. (2009). Fecundity of the autumn spawning herring in the Gulf of Riga (Baltic Sea): comparison between 1950s-1970s and 2000s. BSSC 2009 ABSTRACT BOOK: 7th Baltic Sea Science Congreess; Tallinn, Estonia; August 17-21, 2009. Ed. Lips, U. Tallinn University of Technology: Infotrükk OÜ, 108−108.
 
Arula, T.; Ojaveer, E.; Ojaveer, H.; Põllupüü, M. (2008). LONG-TERM DYNAMICS OF THE SPRING-SPAWNING HERRING LARVAE (CLUPEA HARENGUS MEMBRAS) IN THE GULF OF RIGA (BALTIC SEA). 32nd Annual Larval Fish Conference: 32nd Annual Larval Fish Conference; Kiel, Germany; 4-7 August 2008.. Ed. Catriona Clemmesen. Germany, Kiel, 56−56.
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ProgramTypeNumberNameProject startProject endPrincipal investigatorInstitutionFinancing
MUULLTMI20093 (hange 215079)National work plan for fisheries sector 2020-202101.01.202010.02.2022Redik EschbaumUniversity of Tartu, Faculty of Science and Technology, Estonian Marine Institute1 792 166,00 EUR
MUUSLTMI17556TImpact of trawlers of different sizes on the ecosystem and fish resources of the Gulf of Riga27.12.201715.02.2021Markus VetemaaUniversity of Tartu, Faculty of Science and Technology, Estonian Marine Institute373 400,00 EUR
MEMRUPMEMRUP46Exposure assessment of mercury in different population groups01.01.201823.12.2019Mart SimmUniversity of Tartu, Faculty of Science and Technology, Estonian Marine Institute20 000,00 EUR
SFSF0182579s03Investigations on dynamics and regularities of development of ecological subsystems in the North-eastern Baltic, Gulf of Finland and Gulf of Riga01.01.200331.12.2007Evald OjaveerUniversity of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu350 044,10 EUR
ETFETF8747Ecology and dynamics of various life history stages of selected marine fish populations at different ecosystems regimes01.01.201131.12.2014Henn OjaveerUniversity of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu48 000,00 EUR
SFSF0180005s10From past to future - development of the Baltic Sea populations and ecosystems under dynamic external forcings01.01.201031.12.2015Henn OjaveerUniversity of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu841 560,50 EUR
1.1.7.2.1SLTMI20351 (17496)Spatial and temporal patterns in Pärnu Bay fish composition01.06.202031.05.2022Timo ArulaUniversity of Tartu, Faculty of Science and Technology, Estonian Marine Institute17 000,00 EUR
MUULLTMI16047Studies in commercial fish larvae in Pärnu Bay and the Gulf of Riga01.02.201605.12.2016Timo ArulaUniversity of Tartu, Faculty of Science and Technology, Estonian Marine Institute18 333,34 EUR
MUULLOMI13024Studies in commercial fish larvae in Pärnu Bay and the Gulf of Riga03.04.201302.12.2013Timo ArulaUniversity of Tartu; University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu22 000,00 EUR
MUULLOMI13024/1Studies in commercial fish larvae in Pärnu Bay and the Gulf of Riga10.06.201402.12.2014Timo ArulaUniversity of Tartu; University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu18 333,33 EUR
MUULLOMI13024/2Studies in commercial fish larvae in Pärnu Bay and the Gulf of Riga 201501.03.201504.12.2015Timo ArulaUniversity of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu18 333,33 EUR
MUUSLOMI13115Developing tool for separation of the Baltic herring (Clupea harengus membras) populations in Estonian waters (to be applied in fisheries monitoring)01.08.201318.06.2015Timo ArulaUniversity of Tartu; University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu60 749,00 EUR
MUUSLOMI11123Insufficiently studied commercial fish in Estonian coastal sea: fall spawning herring in the Gulf of Riga and northern Baltic Sea01.07.201130.06.2013Henn OjaveerUniversity of Tartu; University of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu31 454,00 EUR
ETFETF6751Zooplankton dynamics and its implications to upper trophic levels in the ecosystem of the Gulf of Riga, Baltic Sea01.01.200631.12.2009Henn OjaveerUniversity of Tartu, Faculty of Science and Technology (old), Estonian Marine Institute, University of Tartu; University of Tartu, Faculty of Science and Technology (old)61 876,71 EUR